Below is a list of scientific articles specifically focusing on Omura’s whales, including data from Omura’s whales, or making reference to Bryde’s whales that were later identified as Omura’s whales.  This list is undoubtedly incomplete and will be added to as more references are found and/or published.

Madagascar Omura’s Whale Project papers:

From: Cerchio et al. 2019, Figure 1, Frontiers in Marine Science, doi: 10.3389/fmars.2019.00067

From: Cerchio et al. 2019, Figure 1, Frontiers in Marine Science, doi: 10.3389/fmars.2019.00067

Cerchio S, Yamada TK, Brownell Jr. RL. 2019. Global distribution of Omura’s whales (Balaenoptera omurai) and assessment of range-wide threats. Frontiers in Marine Science 6:67. doi: 10.3389/fmars.2019.00067

Cerchio S, Andrianantenaina B, Zerbini A, Pendleton D, Rasoloarijao T, Cholewiak D. 2018. Residency, feeding ecology, local movements and potential isolation of the Madagascar Omura’s whale (Balaenoptera omurai) population. Paper SC/67B/NH09 presented to the IWC Scientific Committee.

Cerchio S, and Yamada T. 2018. Omura’s whale: Balaenopteridae omurai. In: “Encyclopedia of Marine Mammals”, Würsig B, Thewissen JGM, and Kovacs K (eds), 3rd ed, pp. 656-659, Academic Press, San Diego, CA, USA.

Cerchio S, Andrianantenaina B, Lindsay A, Rekdahl M, Andrianarivelo N, and Raoloarijao T. 2015. Omura’s whales (Balaenoptera omurai) in Northwest Madagascar: ecology, behavior and conservation needs. Royal Society Open Science 2: 150301. DOI:10.1098.

Work on other species related to the Madagascar Project:

Cerchio S, Rasoloarijao T, Cholewiak D. 2018. Acoustic monitoring of Blue Whales (Balaenoptera musculus) and other baleen whales in the Mozambique Channel off the Northwest Coast of Madagascar. Progress Report SC/67B/SH/14 presented to IWC Scientific Committee.

Cerchio S, Willson A, Muirhead C, Al Harthi S, Baldwin R, Cholewiak D, Collins T, Minton G, Rasoloarijao T, Sarrouf Willson M. 2018. A new baleen whale song type described for the Western Indian Ocean off Oman and northwest Madagascar. Paper SC/67B/SH24 presented to the IWC Scientific Committee.

Cerchio S, Andrianantenaina B, Guillemaind’Echon T, and Guillemaind’Echon A. 2016. Evidence for the presence of “pygmy” blue whales (Balaenoptera musculus brevicauda) in the Mozambique Channel off the Northwest coast of Madagascar. Paper SC/66b/SH33 presented to IWC Scientific Committee.

Papers related to or mentioning Omura’s whales from other regions:


Kim JH, Kim HW, Kim EM, and Sohn H. 2018. First Record of the Omura’s whale (Balaenoptera omurai) in Korean Waters. Anim. Syst. Evol. Divers. 34, 162-167.

Brownell RL, Jr., de Vos A. and Ilangakoon AD. 2017. Large whale strandings from Sri Lanka between 1889 and 2014. Paper SC/67A/HIM/11 presented to the International Whaling Commission Scientific Committee. 13 pp.

deVos A. 2017. First record of Omura’s whale, Balaenoptera omurai, in Sri Lankan waters. Marine Biodiversity Records 10, 18.

Egami T, and Kojima K. 2017. Records of whaling and societal change in Lamalera, Indonesia. Presentation at 28th Annual meeting of The Cetology Study Group of Japan, Sapporo, Japan, 24-25 June 2017.

Erbe C, Dunlop R, Jenner KCS, Jenner MN, McCauley RD, Parnum I, Parsons M, Rogers T, and Salgado-Kent C. 2b017. Review of Underwater and In-Air Sounds Emitted by Australian and Antarctic Marine Mammals. Acoustics Australia, 45, 179-241.

Notarbartolo di Sciara G, Kerem D, Smeenk C, Rudolph P, Cesario A, Costa M, Elasar M, Feingold D, Fumagalli M, Goffman O, Hadar N, Mebrathu YT, and Scheinin A. 2017. Cetaceans of the Red Sea. CMS Technical Series 33, 86 pp.

Slater, GJ, Goldbogen JA, and Pyenson ND. 2017. Independent evolution of baleen whale gigantism linked to Plio-Pleistocene ocean dynamics. Proc. R. Soc. B, 284, 20170546.

Xu, M., Wang, X., Miao, X., Wu, F., Ma, M., Tao, C., and Zhu, Q. (2017).A stranding of Omura’s whale (Balaenoptera omurai Wada, Oishi and Yamada, 2003) in the Taiwan Strait, China. Aquatic Mammals 43, 289-298. DOI:10.1578/AM.43.3.2017.289

Cypriano‐Souza AL, Meirelles, ACO., Carvalho VL, and Bonatto SL. 2016. Rare or cryptic? The first report of an Omura's whale (Balaenoptera omurai) in the South Atlantic Ocean. Marine Mammal Science, 33, 80-95.

Kym O, Coughran D, Gall M, Irvine L, and Byrne M. 2016. Stranding of Omura’s Whale (Balaenoptera Omurai) in Western Australia. Aquatic Mammals 42, 193-197. DOI 10.1578/AM.42.2.2016.193

Ranjbar S, Dakhteh, SMH, Van Waerebeek K. 2016. Omura's whale Balaenoptera omurai stranding on Qeshm Island, Iran, Persian Gulf: further evidence for a wide (sub)tropical distribution. J. Mar. Biol. Oceanogr. 5, 3. DOI: 10.4172/2324-8661.1000161


Acebes, J.M.V. (2013). Hunting “big fish”: a marine environmental history of a contested fishery in the Bohol Sea. Doctoral dissertation, Perth, Murdoch University.395 pp.

Acebes, J.M.V. (2014). “A history of whaling in the Philippines: a glimpse of the past and current distribution of whales,” in: Historical Perspectives of Fisheries Exploitation in the Indo-Pacific, eds. Christensen J., and Tull M., (Dordrecht: Springer), 83-105.Adulyanukosol, Kanjana, SurasakThongsukdee, SurachaiPassada, TheerawatPrempree, and TanetWannarangsee. 2012. Bryde’s Whales in Thailand. Bangkok: Aksornthai Printing Co.

Garrigue, C, and M Poupon. 2013. Guide D’identification: MammifèresMarins de Nouvelle-Calédonie. Nouméa: Artypo.

Jung, J L, W C Mullié, K Van Waerebeek, M MWagne, A S O Bilal, Z A O Sidaty, L Toomey, E Méheust, and F Marret. 2015. Omura’s Whale off West Africa: Autochthonous Population or Inter-Oceanic Vagrant in the Atlantic Ocean? Marine Biology Research 12, 66-75.

Kershaw, F, M S Leslie, T Collins, R M Mansur, B D Smith, M Gianna, R Baldwin, et al. 2013. “Population Differentiation of 2 Forms of Bryde’s Whales in the Indian and Pacific Oceans.” Journal of Heredity 104 (6): 755–64. /citations?view_op=view_citation&continue=/scholar%3Fhl%3Den%26as_sdt%3D1,5%26scilib%3D1&citilm=1&citation_for_view=EPm-87YAAAAJ:u5HHmVD_uO8C&hl=en&oi=p.

Marx, F G, and R E Fordyce. 2015. “Baleen Boom and Bust: A Synthesis of Mysticete Phylogeny, Diversity and Disparity.” Royal Society Open Science 2 (4): 140434.

Oremus, M., Leqata, J., Hurutarau, J., Taei, S., Donoghue, M., Thompson, K., and Baker, C.S. (2011). Solomon Islands Dolphin Project: Progress report on data collection and analyses. Technical report to the South Pacific Whale research Consortium. 27 pp.

Ponnampalam, L S. 2012.“Opportunistic Observations on the Distribution of Cetaceans in the Malaysian South China, Sulu and Sulawesi Seas and an Updated Checklist of Marine.” Raffles B Zool 60 (1): 221–31.

Rosel, P.E., and Wilcox, L.A. (2014). Genetic evidence reveals a unique lineage of Bryde’s whales in the northern Gulf of Mexico. Endangered Species Research, 25, 19-34.

Rosel, P E, and L A Wilcox. 2014. “Genetic Evidence Reveals a Unique Lineage of Bryde’s Whales in the Northern Gulf of Mexico.” Endangered Species Research 25 (19-34). /citations?view_op=view_citation&continue=/scholar%3Fhl%3Den%26as_sdt%3D1,5%26scilib%3D1%26scioq%3Dbryde%2Bauthor:PE%2Bauthor:rosel&citilm=1&citation_for_view=EPm-87YAAAAJ:u-x6o8ySG0sC&hl=en&oi=p.

Sousa, A.G. and Harris, D. (2015). Description and seasonal detection of two potential whale calls recorded in the Indian Ocean. The Journal of the Acoustical Society of America, 138, 1379-1388.

Van Canneyt, O, G Dorémus, S Laran, V Ridoux, and P Watremez. 2015. “REMMOA Nouvelle-Calédonie Wallis et Futuna: Rapport de Campagne.” Rapport Intermédiaire Pour l’Agence Des Aires Marines Protégées, 65pp.



Acebes, J.M.V. (2009). Historical whaling in the Philippines: origins of ‘indigenous subsistence whaling’, mapping whaling grounds and comparison with current known distribution: a HMAP Asia Project Paper. History of Marine Animal Populations working paper 161.Murdoch University, Perth.37 pp.

Aragones, L.V., Roque, M.A.A., Flores, M.B., Encomienda, R.P., Laule, G.E., Espinos, B.G., ... and Braun, R.C. (2010). The Philippine marine mammal strandings from 1998 to 2009: animals in the Philippines in peril? Aquatic Mammals, 36, 219.

Best, P B. 2001.“Distribution and Population Separation of Bryde’s Whale Balaenoptera edeni off Southern Africa.”Marine Ecology Progress Series. Progress Series 220: 277–89.

Best, P B. 2007.Whales and Dolphins of the Southern African Subregion. Cambridge: Cambridge University Press.

Ilangakoon, A. 2002.Whales and dolphins of Sri Lanka. Colombo: WHT Publications Limited.

Kahn, B. (2001). Komodo National Park Cetacean Surveys: A rapid ecological assessment of cetacean diversity, abundance and distribution. Report to The Nature Conservancy Indonesia Coastal and Marine Program. April 2001.

Kahn, B. (2005). Indonesia Oceanic Cetacean Program Activity Report: January-February 2005. Report to The Nature Conservancy SE Asia Center for Marine Protected Areas. April 2005.

Kahn, B., Wawandono, N.B., and Subijanto, J. (2001).Protecting the cetaceans of Komodo National Park, Indonesia: Positive identification of the rare pygmy Bryde’s whale (Balaenoptera edeni) with the assistance of genetic profiling.Status report to The Nature Conservancy.

LeDuc, R, and A E Dizon. 2002. “Reconstructing the Rorquals Phylogeny: With Comments on the Use of Molecular and Morphological Data for Systematic Study.” In Molecular and Cell Biology of Marine Mammals, edited by CJ Pfeiffer and Paul E. Nachtigall, 100–110. Florida: Krieger.

Ma, M., Zhu, Q., Li, X., Sun, Y., Liu, Y.Y., Liu, Y.N., and Tao, C.H. (2007).Identification of Balaenoptera omurai from an unknown whale vertebra based on Cyt b gene sequence [in Chinese].ActaTheriologicaSinica 27, 288–292.

Mustika, P.T.K. (2006). Marine Mammals in the Savu Sea (Indonesia): Indigenous Knowledge, Threat Analysis and Management Options. MSc. Thesis.James Cook University.231 pp.

Perrin, W. F. 2007. The Philippine fishery for Bryde’s whales in the western North Pacific, 1983-1986. Paper SC/58/RMP1 presented to the International Whaling Commission Scientific Committee.

Reeves, R R, W F Perrin, B L Taylor, C S Baker, and S L Mesnick. 2004. “Report of the Workshop on Shortcomings of Cetacean Taxonomy in Relation to Needs of Conservation and Management.” Edited by R R Reeves, W F Perrin, B L Taylor, C S Baker, and S L Mesnick. NOAA Technical Memorandum, NOAA-NMFS-SWFSC-363:59-60. et al_2004_Fisheries Science_Report of the Workshop on Shortcomings of Cetacean Taxonomy in Relation to Needs of Conservation and Management La Jolla , California Report of the Workshop on Shortcomings of Cetacean.

Sasaki, Takeshi, Masato Nikaido, Shiro Wada, Tadasu K Yamada, Ying Cao, Masami Hasegawa, and Norihiro Okada. 2006. “Balaenoptera Omurai Is a Newly Discovered Baleen Whale That Represents an Ancient Evolutionary Lineage.” Molecular Phylogenetics and Evolution 41 (1): 40–52. doi:10.1016/j.ympev.2006.03.032.

Wada, S, M Oishi, and T K Yamada. 2003. “A Newly Discovered Species of Living Baleen Whale.” Nature 426: 278–81.

Wang, H., Fan, Z., Shen, H., and Peng, Y. 2006. Description of new record species of whales from Chinese coastal waters [in Chinese].  Fisheries Science, 25(2), 85-87. DOI:10.3969/j.issn.1003-1111.2006.02.010

Yamada, T.K. (2009). “Omura’s whale, Balaenoptera omurai”, in The Wild Mammals of Japan, eds. S. Ohdachi, Y. Ishibashi, M. Iwasa, and T. Saitoh (Kyoto: Shoukahoh Book Sellers and Mammalogical Society of Japan), 330–331.

Yamada, T K, L S Chou, S Chantrapornsyl, K Adulyanukosol, S K Chakravarti, M Oishi, S Wada, et al. 2006. “Middle-Sized Balaenopterid Whale Specimens (Cetacea: Balaenopteridae) Preserved at Several Institutions in Taiwan, Thailand, and India.” Memoirs of the National Science Museum, Tokyo 44: 1–10.

Yamada, T K, T Kakuda, and Y Tajima. 2008. “Middle Sized Balaenopterid Whale Specimens in the Philippines and Indonesia.” Memoirs of the National Science Museum, Tokyo 45: 75–83.

Yamada, T K, C Kemper, Y Tajima, AUmetani, H Janetzki, and D Pemberton. 2006. “Marine Mammal Collections in Australia.” National Science Museum Monographs 34: 117–26.

Yang, G, H Liu, K Zhou, and G Ji. 2002. “Identification of a Balaenoptera edeni Specimen by Using Mitochondrial DNA Sequences.” Chinese Journal of Zoology 37 (4): 35–38.



Dolar, M.L., Leatherwood, S., Wood, C.J., Alava, M.N., Hill, C.L., and Aragones, L.V. (1994). Directed fisheries for cetaceans in the Philippines.Reports of the International Whaling Commission 44, 439-49.

Perrin, W F, M L LDolar, and E Ortega. 1996. “Osteological Comparison of Bryde’s Whales from the Philippines with Specimens from Other Regions.” Report SC/47/NP2 to the Scientific Committee of the International Whaling Commission.

Wada, S, and K Numachi. 1991. “Allozyme Analyses of Genetic Differentiation among the Populations and Species of the Balaenoptera.” Reports of the International Whaling Commission, Special Issue 13: 125–1545.

Yoshida, H, and H Kato. 1999. “Phylogenetic Relationships of Bryde’s Whale in the Western North Pacific and Adjacent Waters Inferred from Mitochondrial DNA Sequences.” Marine Mammal Science 15: 1269–86.



 None found.


Kawamura, A. 1977. “On the Food of Bryde’s Whales Caught in the South Pacific and Indian Oceans.” Scientific Reports of the Whales Research Institute 29: 49–58.

Kawamura, A. 1980. “Food Habits of the Bryde’s Whales Taken in the South Pacific and Indian Oceans.” Scientific Reports of the Whales Research Institute 32: 1–23.

Ohsumi, S. (1980). Population study of the Bryde’s whale in the southern hemisphere under scientific permit in the three seasons 1976/77–1978/79. Rep. Int. Whal. Comm. 30, 319-331.

Ohsumi, S. (1978). Provisional report on the Brydes whales caught under special permit in the Southern Hemisphere. Report of the International Whaling Commission, 28, 281-287.

Omura, H., (1977). Review of the occurrence of Bryde’s whale in the northwest Pacific. Rep. Int. Whal. Comm. Spec. Issue 1, 88-91.


Before 1970

Anonymous (1955).Hong Kong's whale. Spectrum: The Magazine of the Hong Kong University Science Society 4: 33-41.

Omura, H., (1962). Further information on Bryde’s whale from the coast of Japan. Scientific reports of the Whales Research Institute, 16, 7-18.